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The Wings of Pegasus are one of the Three Sacred Treasures. They grant Pit the ability of full, unlimited flight. They're only unsealed for the final encounter in the Palace in the Sky. A lesser version, known as an Angel's Feather, grants Pit temporary flight as a recovery method.
The Wings of Pegasus appear in the Palace in the Sky in Pit's final battle with Medusa. In the American NES version, players had to simply direct which way the wings would take him (though the screen was always automatically scrolling to the right), but in the Japanese version, players had to repeatedly hit the jump button in order to keep Pit flying. These wings will never exhaust, though are still only usable in the last level.
These wings give Pit flight, but the jump button must be continuously pressed to keep him in the air.
All of the Three Sacred Treasures appear in this game, though Palutena disables the Wings of Pegasus because she fears they will not function properly after so many years in storage, forcing Pit to rely on the time-limited Power of Flight instead. However, Pit uses the Wings of Pegasus in the fight against Medusa, as the Power of Flight has already been used. They create a magenta pink glow when activated.